:root {
  color-scheme: light;
  --bg: #f6f7f4;
  --surface: #ffffff;
  --text: #1f2a24;
  --muted: #5d6a62;
  --line: #d9ded8;
  --accent: #365f4b;
  --accent-strong: #234536;
}

* {
  box-sizing: border-box;
}

body {
  margin: 0;
  background: var(--bg);
  color: var(--text);
  font-family: system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  line-height: 1.5;
}

a {
  color: inherit;
}

button,
input {
  font: inherit;
}

.site-header {
  align-items: center;
  background: var(--surface);
  border-bottom: 1px solid var(--line);
  display: flex;
  gap: 24px;
  justify-content: space-between;
  min-height: 64px;
  padding: 0 32px;
}

.brand {
  color: var(--accent-strong);
  font-weight: 700;
  text-decoration: none;
}

.top-nav,
.top-nav form {
  align-items: center;
  display: flex;
  gap: 12px;
}

.top-nav {
  flex-wrap: wrap;
  justify-content: flex-end;
  min-width: 0;
}

.top-nav a,
.top-nav span {
  white-space: nowrap;
}

.page-shell {
  margin: 0 auto;
  max-width: 960px;
  padding: 40px 24px;
}

.panel,
.auth-panel {
  background: var(--surface);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 24px;
}

.auth-panel {
  margin: 48px auto;
  max-width: 420px;
}

.auth-panel form {
  display: grid;
  gap: 12px;
}

input {
  border: 1px solid var(--line);
  border-radius: 6px;
  padding: 10px 12px;
  width: 100%;
}

button {
  background: var(--accent);
  border: 0;
  border-radius: 6px;
  color: #ffffff;
  cursor: pointer;
  padding: 10px 14px;
}

button:hover {
  background: var(--accent-strong);
}

.panel + .panel {
  margin-top: 24px;
}

.panel h1,
.panel h2 {
  margin-top: 0;
}

.exam-answer-block {
  border-bottom: 1px solid var(--line);
  padding: 18px 0;
}

.exam-answer-block:first-of-type {
  padding-top: 0;
}

.exam-answer-block:last-of-type {
  border-bottom: 0;
  padding-bottom: 0;
}

.data-table {
  border-collapse: collapse;
  width: 100%;
}

.data-table th,
.data-table td {
  border-bottom: 1px solid var(--line);
  padding: 8px 10px;
  text-align: left;
  vertical-align: top;
}

.data-table th {
  color: var(--muted);
  font-weight: 600;
}

.data-table:focus-visible {
  outline: 2px solid var(--accent);
  outline-offset: 2px;
}

.form-grid {
  display: grid;
  gap: 12px;
  max-width: 480px;
}

.form-grid label {
  font-weight: 600;
}

.form-grid select,
.form-grid textarea {
  border: 1px solid var(--line);
  border-radius: 6px;
  padding: 10px 12px;
  width: 100%;
}

.actions {
  align-items: center;
  display: flex;
  flex-wrap: wrap;
  gap: 12px;
  margin: 16px 0;
}

.button-link {
  background: var(--accent);
  border-radius: 6px;
  color: #ffffff;
  display: inline-block;
  padding: 10px 14px;
  text-decoration: none;
}

.button-link:hover {
  background: var(--accent-strong);
}

.button-secondary {
  background: transparent;
  border: 1px solid var(--accent);
  color: var(--accent-strong);
}

.button-secondary:hover {
  background: var(--bg);
}

.badge {
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 999px;
  display: inline-block;
  font-size: 0.85em;
  padding: 2px 10px;
  white-space: nowrap;
}

.badge-error {
  background: #fbeaea;
  border-color: #d9a0a0;
  color: #8a2525;
}

.badge-warning {
  background: #fdf6e3;
  border-color: #d9c48a;
  color: #7a5d12;
}

.badge-ok {
  background: #e9f3ec;
  border-color: #9fc4ab;
  color: #234536;
}

.messages {
  display: grid;
  gap: 8px;
  list-style: none;
  margin: 0 0 24px;
  padding: 0;
}

.messages li {
  background: #e9f3ec;
  border: 1px solid #9fc4ab;
  border-radius: 6px;
  padding: 10px 14px;
}

.messages li.error {
  background: #fbeaea;
  border-color: #d9a0a0;
}

.errorlist {
  color: #8a2525;
  list-style: none;
  margin: 0;
  padding: 0;
}

.muted {
  color: var(--muted);
}

.eyebrow {
  color: var(--muted);
  font-size: 0.78rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  margin: 0 0 6px;
  text-transform: uppercase;
}

.species-backlink {
  margin-top: 0;
}

.species-header {
  align-items: center;
  display: flex;
  gap: 18px;
  justify-content: space-between;
}

.species-header h1,
.species-list-section h2,
.species-section h2,
.species-facts h2 {
  margin-top: 0;
}

.species-latin {
  color: var(--muted);
  font-family: Georgia, "Times New Roman", serif;
  font-style: italic;
  margin: 4px 0 0;
}

.species-image-grid {
  display: grid;
  gap: 20px;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  margin: 24px 0;
}

.species-image-card {
  margin: 0;
  padding: 14px;
}

.species-image-card img {
  aspect-ratio: 4 / 3;
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 6px;
  display: block;
  object-fit: cover;
  width: 100%;
}

.species-image-card figcaption {
  display: grid;
  gap: 2px;
  margin-top: 10px;
}

.species-image-card figcaption span {
  color: var(--muted);
  font-size: 0.9rem;
}

.species-detail-layout {
  align-items: start;
  display: grid;
  gap: 24px;
  grid-template-columns: minmax(0, 1fr) 340px;
  margin-bottom: 40px;
}

.species-text-stack {
  display: grid;
  gap: 20px;
}

.species-section p {
  line-height: 1.65;
  margin: 0;
}

.species-facts {
  background: #fbf7e9;
  border-color: #d9c48a;
}

.species-facts h2 {
  border-bottom: 1px solid #d9c48a;
  color: #6f5512;
  font-size: 1.05rem;
  padding-bottom: 10px;
}

.species-facts dl,
.species-summary-facts {
  display: grid;
  gap: 0;
  margin: 0;
}

.species-facts dl div,
.species-summary-facts div {
  border-bottom: 1px solid rgba(217, 196, 138, 0.45);
  display: grid;
  gap: 6px;
  grid-template-columns: 120px minmax(0, 1fr);
  padding: 10px 0;
}

.species-facts dl div:last-child,
.species-summary-facts div:last-child {
  border-bottom: 0;
}

.species-facts dt,
.species-summary-facts dt {
  color: #6f5512;
  font-weight: 700;
}

.species-facts dd,
.species-summary-facts dd {
  margin: 0;
}

.species-list-section {
  margin-top: 38px;
}

.species-card-grid {
  margin-top: 18px;
}

.species-summary-card {
  justify-content: space-between;
}

.species-summary-head {
  display: block;
  margin-bottom: 12px;
  min-height: 44px;
  position: relative;
}

.species-summary-image {
  aspect-ratio: 4 / 3;
  background: #f6f2ec;
  border: 1px solid var(--line);
  border-radius: 6px;
  display: block;
  height: auto;
  max-height: 190px;
  object-fit: cover;
  width: 100%;
}

.species-summary-head .badge {
  background: rgba(255, 255, 255, 0.94);
  position: absolute;
  right: 10px;
  top: 10px;
  white-space: nowrap;
}

.species-summary-card h3 {
  margin: 0 0 4px;
}

.species-summary-facts {
  background: var(--bg);
  border-radius: 6px;
  font-size: 0.9rem;
  margin-top: 16px;
  padding: 4px 12px;
}

.species-summary-facts div {
  grid-template-columns: 88px minmax(0, 1fr);
}

.species-card-action {
  margin-top: 20px;
  text-align: center;
  width: 100%;
}

.reference-hero {
  align-items: end;
  background: var(--surface);
  border: 1px solid var(--line);
  border-radius: 8px;
  display: grid;
  gap: 24px;
  grid-template-columns: minmax(0, 1fr) auto;
  padding: 28px;
}

.reference-hero h1 {
  margin: 0;
}

.reference-hero p:not(.eyebrow) {
  color: var(--muted);
  font-size: 1.05rem;
  margin: 10px 0 0;
  max-width: 680px;
}

.reference-stats {
  display: grid;
  gap: 10px;
  grid-template-columns: repeat(3, minmax(74px, 1fr));
  margin: 0;
}

.reference-stats div {
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 6px;
  padding: 10px 12px;
  text-align: center;
}

.reference-stats dt {
  color: var(--muted);
  font-size: 0.78rem;
  font-weight: 700;
  text-transform: uppercase;
}

.reference-stats dd {
  color: var(--accent-strong);
  font-size: 1.4rem;
  font-weight: 800;
  margin: 2px 0 0;
}

.reference-quick-grid {
  display: grid;
  gap: 16px;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  margin-top: 20px;
}

.reference-tile {
  background: var(--surface);
  border: 1px solid var(--line);
  border-radius: 8px;
  color: var(--text);
  display: grid;
  gap: 6px;
  min-height: 154px;
  padding: 18px;
  text-decoration: none;
}

.reference-tile:hover {
  border-color: var(--accent);
}

.reference-tile-primary {
  background: #edf4ef;
  border-color: #9fc4ab;
}

.reference-tile-kicker {
  color: var(--muted);
  font-size: 0.76rem;
  font-weight: 800;
  text-transform: uppercase;
}

.reference-tile strong {
  color: var(--accent-strong);
  font-size: 1.05rem;
}

.reference-tile span:last-child {
  color: var(--muted);
  font-size: 0.92rem;
}

.reference-section {
  margin-top: 36px;
}

.reference-section-head h2 {
  margin: 0;
}

.reference-chapter-list {
  display: grid;
  gap: 18px;
  margin-top: 18px;
}

.reference-chapter {
  background: var(--surface);
  border: 1px solid var(--line);
  border-radius: 8px;
  display: grid;
  gap: 22px;
  grid-template-columns: 300px minmax(0, 1fr);
  padding: 22px;
}

.reference-chapter h3 {
  margin: 2px 0 8px;
}

.reference-chapter-meta {
  color: var(--muted);
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
  margin-top: 16px;
}

.reference-chapter-meta span {
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 999px;
  font-size: 0.82rem;
  padding: 3px 9px;
}

.reference-lesson-list {
  counter-reset: reference-lesson;
  display: grid;
  gap: 8px;
  list-style: none;
  margin: 0;
  padding: 0;
}

.reference-lesson-list a {
  align-items: center;
  border: 1px solid var(--line);
  border-radius: 6px;
  color: var(--text);
  display: grid;
  gap: 12px;
  grid-template-columns: 34px minmax(0, 1fr);
  min-height: 50px;
  padding: 8px 12px;
  text-decoration: none;
}

.reference-lesson-list a:hover {
  border-color: var(--accent);
}

.reference-lesson-list span {
  align-items: center;
  background: var(--bg);
  border-radius: 50%;
  color: var(--accent-strong);
  display: inline-flex;
  font-size: 0.86rem;
  font-weight: 800;
  height: 30px;
  justify-content: center;
  width: 30px;
}

.reference-lesson-list strong {
  font-size: 0.96rem;
}

@media (max-width: 780px) {
  .species-header,
  .species-detail-layout,
  .species-image-grid {
    grid-template-columns: 1fr;
  }

  .reference-hero,
  .reference-chapter {
    grid-template-columns: 1fr;
  }

  .reference-quick-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .species-header {
    align-items: flex-start;
    flex-direction: column;
  }
}

@media (max-width: 700px) {
  .site-header {
    align-items: flex-start;
    flex-direction: column;
    gap: 10px;
    padding: 14px 20px;
  }

  .top-nav {
    justify-content: flex-start;
    width: 100%;
  }

  .top-nav form {
    margin: 0;
  }

  .page-shell {
    padding: 24px 16px;
  }

  .panel,
  .auth-panel,
  .lesson-content,
  .lesson-metadata-panel {
    padding: 20px;
  }

  .actions {
    align-items: stretch;
  }

  .actions > *,
  .button-link,
  button {
    max-width: 100%;
    white-space: normal;
  }

  .data-table {
    display: block;
    overflow-x: auto;
  }

  .reference-stats,
  .reference-quick-grid {
    grid-template-columns: 1fr;
  }

  .reference-hero {
    padding: 22px;
  }
}

/* Learning Modules & Curriculum Styles */
.modules-grid {
  display: grid;
  gap: 20px;
  grid-template-columns: repeat(auto-fill, minmax(280px, 1fr));
  margin-top: 20px;
}

.module-card {
  display: flex;
  flex-direction: column;
  justify-content: space-between;
}

.module-meta {
  font-size: 0.85em;
  font-weight: 700;
  color: var(--accent);
  text-transform: uppercase;
  letter-spacing: 0.05em;
  margin-bottom: 6px;
}

.progress-container {
  margin-top: 16px;
}

.progress-bar {
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 99px;
  height: 12px;
  margin-top: 6px;
  overflow: hidden;
  position: relative;
  width: 100%;
}

.progress-bar-fill {
  background: var(--accent);
  height: 100%;
  transition: width 0.3s ease;
}

.progress-label {
  display: flex;
  font-size: 0.85em;
  justify-content: space-between;
  color: var(--muted);
}

.lessons-list {
  display: grid;
  gap: 12px;
  list-style: none;
  margin: 20px 0;
  padding: 0;
}

.lesson-item {
  align-items: center;
  background: var(--surface);
  border: 1px solid var(--line);
  border-radius: 8px;
  display: flex;
  gap: 16px;
  justify-content: space-between;
  padding: 16px 20px;
  transition: transform 0.2s, box-shadow 0.2s;
  text-decoration: none;
}

.lesson-item:hover {
  box-shadow: 0 4px 12px rgba(0,0,0,0.05);
  transform: translateY(-2px);
}

.lesson-info {
  display: flex;
  flex-direction: column;
}

.lesson-title {
  font-weight: 600;
}

.lesson-content {
  background: var(--surface);
  border: 1px solid var(--line);
  border-radius: 8px;
  margin-top: 24px;
  padding: 32px;
}

.lesson-content h1,
.lesson-content h2,
.lesson-content h3 {
  color: var(--accent-strong);
  margin-top: 24px;
  margin-bottom: 12px;
}

.lesson-content h1 {
  border-bottom: 2px solid var(--line);
  font-size: 2em;
  padding-bottom: 8px;
}

.lesson-content h2 {
  font-size: 1.5em;
}

.lesson-content h3 {
  font-size: 1.25em;
}

.lesson-content p {
  line-height: 1.6;
  margin-bottom: 16px;
  text-align: justify;
}

.lesson-content ul {
  margin-bottom: 16px;
  padding-left: 24px;
}

.lesson-content li {
  margin-bottom: 6px;
}

.lesson-content img {
  border: 4px solid var(--surface);
  border-radius: 8px;
  box-shadow: 0 8px 24px rgba(0, 0, 0, 0.12);
  display: block;
  margin: 32px auto;
  max-width: 100%;
}

.lesson-content hr {
  border: 0;
  border-top: 1px solid var(--line);
  margin: 32px 0;
}

.lesson-content table {
  border-collapse: collapse;
  margin: 24px 0;
  width: 100%;
}

.lesson-content th,
.lesson-content td {
  border: 1px solid var(--line);
  padding: 10px 12px;
  text-align: left;
}

.lesson-content th {
  background: var(--bg);
  font-weight: 600;
}

.lesson-metadata-panel {
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 8px;
  margin-top: 24px;
  padding: 20px;
}

.lesson-metadata-panel h4 {
  margin: 0 0 10px;
}

.lesson-metadata-panel ul {
  list-style: none;
  margin: 0;
  padding: 0;
}

.lesson-metadata-panel li {
  font-size: 0.9em;
  margin-bottom: 6px;
}

/* Question Practice Styles */
.option-label {
  display: flex;
  align-items: center;
  gap: 12px;
  background: var(--bg);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 14px 20px;
  margin-bottom: 12px;
  cursor: pointer;
  transition: background 0.2s, border-color 0.2s;
}

.option-label:hover {
  background: var(--surface);
  border-color: var(--accent);
}

.option-label input[type="checkbox"],
.option-label input[type="radio"] {
  width: auto;
  margin: 0;
}

.option-result {
  border-radius: 8px;
  padding: 14px 20px;
  margin-bottom: 12px;
  display: flex;
  justify-content: space-between;
  align-items: center;
  border: 1px solid var(--line);
}

.option-result.correct-option {
  background: #e9f3ec;
  border-color: #9fc4ab;
  color: #234536;
}

.option-result.wrong-selected {
  background: #fbeaea;
  border-color: #d9a0a0;
  color: #8a2525;
}

.option-result.correct-selected {
  background: #e9f3ec;
  border-color: #9fc4ab;
  color: #234536;
  font-weight: bold;
}

.explanation-box {
  background: #fdf6e3;
  border: 1px solid #d9c48a;
  border-radius: 8px;
  padding: 20px;
  margin: 24px 0;
  color: #7a5d12;
}

.explanation-box h3 {
  color: #7a5d12;
  margin-top: 0;
  margin-bottom: 8px;
  font-size: 1.1em;
}